Efficient expansion of global protected areas requires simultaneous planning for species and ecosystems

Abstract The Convention on Biological Diversity (CBD)’s strategic plan advocates the use of environmental surrogates, such as ecosystems, as a basis for planning where new protected areas should be placed. However, the efficiency and effectiveness of this ecosystem-based planning approach to adequately capture threatened species in protected area networks is unknown. We tested the application of this approach in Australia according to the nation’s CBD-inspired goals for expansion of the national protected area system. We set targets for ecosystems (10%